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In 1998, in the television drama The Echo, she played Amanda Powell.The next year, she played in the science fiction horror film Event Horizon as Lieutenant Starck, executive officer of the research and rescue ship Lewis and Clark, sent to rescue crew of the long-lost experimental ship Event Horizon.Joely joined the cast of TV series Titanic: Blood and Steel in which she played the role of Countess Markievicz. Richardson had an affair with theatre producer Archie Stirling, resulting in the failure of Stirling's marriage to Diana Rigg in 1990.
The role reunited her with her former husband Tim Bevan, who was part of the show's production team.
She is the niece of actors Lynn Redgrave and Corin Redgrave and cousin of actress Jemma Redgrave, who is five days younger than Richardson.
Joely appeared as an extra at the age of three in the 1968 version of The Charge of the Light Brigade, directed by her father.
Richardson was born in Marylebone, London, to the theatrical Redgrave family, the daughter of actress Vanessa Redgrave and director Tony Richardson, and the granddaughter of actors Sir Michael Redgrave and Rachel Kempson, Lady Redgrave.
Actress Natasha Richardson (1963–2009) was her sister, and through her, she is the sister-in-law of actor Liam Neeson, and the aunt of Micheál and Daniel Neeson.